Apoecus phaedusoides (Thiele, 1931)

Apoecus phaedusoides (Thiele, 1931)

Ena (?Heudiella) phaedusoides was proposed as a replacement name for Buliminus clausiliaeformis Bavay and Dautzenberg 1912 (p. 27-28; pl. 4, figs. 4-6 [images above]) non Möllendorff, 1901 by Thiele (1931: 521; fig. 611). Zilch (1959: 187) placed this taxon in Coccoderma Möllendorff, 1901 [Type Buliminus granulatus Möllendorff, 1901 OD (Schileyko, 1998: 194-5); see image of lectotype from Yen (1939: pl. 7, fig. 14)] non Zittel, 1887 [Pisces: a Coelocanth!], which is plausible assignment but an unavailable generic name. Schileyko (1998: 212) states this species could be a Pseudonapaeus Westerlund, 1887 [Type Buliminus asiaticus von Martens, 1881 SD Lindholm, 1922; see image from Schileyko (1998: 197; fig. 243A)] or Mirus Albers, 1850 [Type Bulimus cantori Philippi, 1844 monotypy; see image from Yen (1939: pl. 7, fig. 1)]. Once assigned to Heudiella Annandale, 1924 [Type H. oliveriana Annandale, 1924 OD; see Annandale's type figure taken from Schileyko (1998: 212; fig.262) above] from Chekiang, China, that taxon has been considered a juvenile clausiliid (Nordsieck, 2000) and is now sunken into Macrophaedusa Möllendorff, 1883 <https://molluscabase.org/aphia.php?p=taxdetails&id=1375420>; thus Heudiella is not a suitable placement for the taxon under consideration. Only recently has it found a home in Apoecus [Type Buliminus colonus Möllendorff, 1895] according to <https://molluscabase.org/aphia.php?p=taxdetails&id=1375431> (editor Páll-Gergely, 2019), which appears to be supported by Köhler et. al. (2016). The genus is an almost exclusively dextral group with about thirty species spread from the SE Asian mainland into New Guinea. As a final testament to the elusive higher taxonomy of this snail, Schileyko (ibid: 182) placed Apoecus in the Pachnodidae (with a "?") and continued: "It is possible that the genus is an aberrant representative of the Camaenidae."

Bavay, A. and P. Dautzenberg, 1912. Description de coquilles nouvelles de l'Indo-Chine (7e suite). Jour. Conchyliologie LX: 1-54, plates I-VI.
Köhler F., F. Criscione, L. Burghardt, & V. Kessner V., 2016. The Enidae of Timor (Stylommatophora: Orthurethra). Molluscan Research. 37(1): 8-16. 10 August.
Nordsieck, H., 2000. Heudiella oliveriana Annandale 1924 from China is not a buliminid but a juvenile clausiliid (Gastropoda: Stylommatophora: Clausiliidae). Mitteilungen der Deutschen Malakozoologischen Gesellschaft 65: 17-19.
Schileyko, A. A., 1998. Treatise on Recent terrestrial pulmonate molluscs 2. Gastrocoptidae, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachnodidae, Enidae, Sagdidae. Ruthenica supplement 2: (ii) + 129-261 incl. numerous text figs.
Thiele, J. 1929-1935. Handbuch der Systematischen Weichtierkunde. Bd 1. Jena: Gustav Fischer, vi + 778 pp. Erster Teil (1929): 1-376; Zweiter Teil (1931): 377-778. Dritter Teil (1934): 779-1022; Vierter Teil (1935): 1023-1154 + xix.
Yen, T.-C., 1939. Die chinesiischen Land- und Süsswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 444: 1-223. July 31.
Zilch, A., 1959-1960.  Gastropoda von Wilhelm Wenz. Teil 2 Euthyneura. Gebrüder Borntraeger, Berlin-Nikolassee, xii + pp. 1-834. pp. 1-200 17 July, 1959; pp. 201-400 25 November, 1959; pp. 401-600 30 March, 1960; pp. 601-835 15 August, 1960.

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